DARWIN Y LA PARADOJA DE LAS ISLAS VACIAS
2009
Blanco Juan Felipe
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3.0cm 70.85pt 3.0cm; mso-header-margin:36.0pt; mso-footer-margin:36.0pt; mso-paper-source:0;} div.Section1 {page:Section1;} --> <!--[if gte mso 10]> <style> /* Style Definitions */ table.MsoNormalTable {mso-style-name:"Tabla normal"; mso-tstyle-rowband-size:0; mso-tstyle-colband-size:0; mso-style-noshow:yes; mso-style-priority:99; mso-style-qformat:yes; mso-style-parent:""; mso-padding-alt:0cm 5.4pt 0cm 5.4pt; mso-para-margin:0cm; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:11.0pt; font-family:"Calibri","sans-serif"; mso-ascii-font-family:Calibri; mso-ascii-theme-font:minor-latin; mso-fareast-font-family:"Times New Roman"; mso-fareast-theme-font:minor-fareast; mso-hansi-font-family:Calibri; mso-hansi-theme-font:minor-latin; mso-bidi-font-family:"Times New Roman"; mso-bidi-theme-font:minor-bidi;} </style> <![endif]--> <p class="MsoNormal" style="margin-bottom: 0.0001pt; text-align: center; line-height: 150%;" align="center"><strong><span style="font-size: 12pt; line-height: 150%; font-family: ";Arial";,";sans-serif";;">RESUMEN</span></strong></p> <p class="MsoNormal" style="margin-bottom: 0.0001pt; text-align: justify; line-height: 150%;"><span style="font-size: 12pt; line-height: 150%; font-family: ";Arial";,";sans-serif";;">Aunque se conoce el interés y fascinación de Darwin por la naturaleza marina a través de sus tratados sobre arrecifes coralinos, islas oceánicas y balanos, poco se comenta sobre los múltiples ejemplos dulceacuícolas que él utilizó para plantear la “Teoría de la Migración” dentro de “El Origen”. Esta plantea que la fauna y la flora se “dispersa” desde los continentes hacia las islas oceánicas. Sin embargo, en la islas muy alejadas de los centros de origen la probabilidad de colonización es muy baja y por lo tanto solo se encuentra una fracción de la biota continental, pudiendo considerarse en algunos casos “vacías”. Aunque las corrientes de agua de dichas islas<span> </span>también se consideran “vacías”, Darwin propuso un mecanismo para explicar la presencia peces “dulceacuícolas” en las mismas, y por lo tanto no podrían considerarse “vacías”, lo cual plantea una paradoja. El mecanismo por él planteado coincide con el ciclo de vida conocido actualmente como diádromo en el cual los individuos de muchas especies de peces, camarones y gasterópodos requieren de aguas marinas y dulces para completar su desarrollo. La diadromía es una convergencia evolutiva que apareció en diferentes linajes a partir de ancestros tanto marinos como dulceacuícolas. En este ensayo se discute la evolución de la fauna dulceacuícola insular, sus implicaciones ecológicas y evolutivas, y algunos modelos experimentales. Finalmente, se discute sobre los impactos de las represas en islas tropicales y subtropicales en las especies diádromas.</span></p> <p class="MsoNormal" style="margin-bottom: 0.0001pt; text-align: justify; line-height: 150%;"><span style="font-size: 12pt; line-height: 150%; font-family: ";Arial";,";sans-serif";;"> </span></p> <p class="MsoNormal" style="margin-bottom: 0.0001pt; text-align: justify; line-height: 150%;"><strong><span style="font-size: 12pt; line-height: 150%; font-family: ";Arial";,";sans-serif";;">Palabras clave:</span></strong><span style="font-size: 12pt; line-height: 150%; font-family: ";Arial";,";sans-serif";;"> Teoría de la migración, biogeografía insular, diadromía, quebradas insulares, El Origen de las Especies, Charles Darwin.<strong></strong></span></p> <p class="MsoNormal" style="margin-bottom: 0.0001pt; text-align: justify; line-height: 150%;"><span style="font-size: 12pt; line-height: 150%; font-family: ";Arial";,";sans-serif";;"> </span></p> <p class="MsoNormal" style="margin-bottom: 0.0001pt; text-align: center; line-height: 150%;" align="center"><strong><span style="font-size: 12pt; line-height: 150%; font-family: ";Arial";,";sans-serif";;">ABSTRACT</span></strong></p> <p class="MsoNormal" style="margin-bottom: 0.0001pt; text-align: justify; line-height: 150%;"><span style="font-size: 12pt; line-height: 150%; font-family: ";Arial";,";sans-serif";;" lang="EN-US">Although Darwin’s fascination and interest on marine nature are well known due to his treatises about coral reefs, oceanic islands, and barnacles, little is commented about the many “freshwater” examples that he provided for supporting a “The Migration Theory” in his book “The Origin of the Species”. According to this theory, faunas and floras disperse from the continents towards oceanic islands. However, those islands located at great distance from the center of origin are very unlikely to be colonized by immigrants, and therefore only a fraction of the continental biota is represented in these islands, and they could be considered as empty. Streams in these islands could also be as empty, however, Darwin stated a mechanism to explain the presence of freshwater fish in these streams, unaware of an emerging paradox. This mechanism is consistent with the current definition of a diadromous life cycle observed in fish, shrimp and snails that rely on migrations between marine and freshwaters to complete their development. This life cycle is an evolutionary convergence among unrelated lineages that evolved from both marine and freshwater ancestors. This essay discusses the evolution of insular freshwater faunas, their implications as ecological and evolutionary forces, and some experimental models. A final discussion presented on the impacts of dams on diadromous fauna in tropical and subtropical islands.</span></p> <p class="MsoNormal" style="margin-bottom: 0.0001pt; text-align: justify; line-height: 150%;"><span style="font-size: 12pt; line-height: 150%; font-family: ";Arial";,";sans-serif";;"> </span></p> <p class="MsoNormal" style="margin-bottom: 0.0001pt; text-align: justify; line-height: 150%;"><strong><span style="font-size: 12pt; line-height: 150%; font-family: ";Arial";,";sans-serif";;" lang="EN-US">Key words:</span></strong><span style="font-size: 12pt; line-height: 150%; font-family: ";Arial";,";sans-serif";;" lang="EN-US"> Migration Theory, island biogeography, diadromy, island streams, The Origin of the Species, Charles Darwin.</span></p>
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