The Quaternary origins of the modern British mammal fauna

The origins of the modern British mammal fauna as a recognizable assemblage are traceable through preceding temperate woodland phases at least as far back as the early part of the Middle Pleistocene. Earlier Quaternary mammal assemblages have more in common with those of the Pliocene than those of the Middle Pleistocene, but evidence for gaps in the British Early Pleistocene sequence precludes inferences concerning the nature of this change in faunal composition. A major difficulty in reconstructing Quaternary faunal histories is in establishing stable correlations between the fragmentary terrestrial sequences from which the fossil record has been recovered and in assembling those sequences into chronological order. Existing reconstructions of Quaternary events in Britain are of hybrid origin, a mix of palynological biostratigraphy based on assumed monocyclic floral successions for a small number of major interglacial stages, with intervening cold stages identified by characteristic glacial or periglacial lithologies and structures. While this framework has served as a useful basis for some Quaternary disciplines, some interpretations of the mammalian evidence do not fit it well. Attention is given here to information derived from fossil mammal assemblages which may be used to help determine the number and sequence of Middle and Late Pleistocene temperate woodland phases with faunas showing close links with that of the Flandrian. While much of this information is not new, it has been largely ignored in the construction of some of the more widely known schemes for Quaternary subdivision. At present five such episodes can be identified and characterized prior to the present interglacial, though further refinement may br possible as work on a more detailed account of key sites arid their palaeontology progresses.