Swarming, supersedure and the mating system of a natural population of honey bees (Apis mellifera capensis)

abeille domestique abeja reina accouplement afrique du sud apis mellifera capensis bee colonies behaviour colonia de abejas colonie d'abeilles comportamiento comportement copulation especies essaimage honeybees mating periodicidad periodicity queen bees reine d'abeille reproduction saison seasonal cycle seasons south africa species swarming