New Finds and Ecology of the Rare Liverworts <i>Scapania apiculata</i>, <i>Scapania carinthiaca,</i> and <i>Scapania scapanioides</i> in Austria
Michaela Kropik | Harald G. Zechmeister
<i>Scapania apiculata</i>, <i>Scapania carinthiaca</i>, and <i>Scapania scapanioides</i> are rare deadwood-dwelling liverworts threatened across Europe. <i>Scapania carinthiaca</i> is thus listed in the Habitats Directive. However, their distribution data are scattered, and their ecologic demands are insufficiently studied. Here, we present new locations and data on the ecology of the species, which resulted from a targeted search in selected regions of Austria. We found ten new sites each for <i>Scapania apiculata</i> and <i>Scapania scapanioides</i> and twenty for <i>Scapania carinthiaca</i>. Reproduction was exclusively asexual. The macroclimates of all known locations in Austria did not differ significantly between the three species. It was consistently wet, with a mean annual precipitation of 1615.3 mm, a high evenness of rainfall, and a low desiccation risk. The mean temperature averaged 7.4 °C. The habitat was shaded dead wood of <i>Picea abies</i>, <i>Abies alba</i>, and <i>Fagus sylvatica</i> of all decay stages at a median distance of 2.5 m from streams or springy areas in semi-natural forests of montane and submontane regions. Thus, high deadwood volumes under a suitable climate are a prerequisite for the occurrences of the species. The number of locations of new finds has more than doubled in Austria and thus in Europe.
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