Effect of pre-harvest application of ethylene inhibitors on polygalacturonase, cellulase and malic-dehydrogenase enzyme activities of apple during cold storage

acide malique acido malico almacenamiento annona squamosa apples cellulase celulasa cold enzimas enzyme enzyme inhibitors enzymes ethylene etileno frio froid inhibidores de enzimas inhibiteur d'enzyme inhibition malate dehydrogenase malato deshidrogenasa malic acid manzana poligalacturonasa polygalacturonase pomme stockage storage